The position of the Lublin and Volyn (Volhynian) uplands within the geobotanical zonation system is ambiguous and disputable. An analysis of the syntaxonomical diversity and landscape distribution of the egetation of the Lublin and Volyn uplands revealed their similarity, and we consider this whole territory as the Lublin-Volyn geobotanical district of hornbeam-oak, oak forests, and steppe meadows. This district differs from the adjacent territories and is classified within the Central European mixed forests ecoregion. Typical and diagnostic syntaxa have been identified, and ecological-coenotic profiles have been constructed to characterize the distribution patterns of shrub-forest and herbaceous vegetation within the landscape. Based on calculations of phytosociological indicators for the main ecological factors, graphical patterns of their variation were developed. It was determined that the characteristic forests of the district, Tilieto-Carpinetum (Carpinion betulis), are represented by four variants (T.-C. var. galeobdolosum; T.-C. var. hepaticosum; T.-C. var. caricosum pilosae; T.-C. var. isopyrosum). The district is also characterized by the presence of syntaxa associated with carbonate soils, both of the marsh type (Caricion davallianae) and the meadow-steppe type (Cirsio-Brachypodion, Festucion valesiacae), with the dominance of Carex humilis and Stipa capillata. Using DCA-analysis, the correlation between ecological factor indicators and the distribution of plant communities within the ecological space has been established, forming six fields distributed along three vectors: by moisture (aquatic and marsh communities), xerophytism, and chemical properties of soils (meadow-steppe communities), as well as the structure of coenoses with a specific ombroregime (forest communities).

Keywords: correlation analysis, ecological-coenotic profiles, Lublin-Volyn geobotanical district, synphytoindication, syntaxonomy, topological differentiation, vegetation

Full text: PDF (Ukr) 2.94M

1. Atlas pochv Ukrainskoy SSR. 1979. Eds. N.K. Krupskiy, N.I. Polupan. Kyiv: Urozhay, 68 pp.
2. Bilyk H.I., Bradis Ye.M. 1962. A geobotanical zoning of the Ukrainian SSR. Ukrainian Botanical Journal, 19(4): 23–32.
3. Bohutskyi A.B. 1998. Lesovyi pokryv Volynskoi vysochyny. In: Ukrainske Polissya: vchora, sohodni, zavtra. Eds. A.B. Bohutskyi, O.A. Bohutskyi, P.K. Voloshyn. Lutsk: Nadstyr’ya, pp. 105–107.
4. Bohutskyi A., Holub B., Lanchont M. 2007. Volynska vysochyna: holovni rysy heolohichnoi budovy ta relyefu. In: Problemy serednopleystotsenovoho interhlyatsialu: materialy XIV ukrainsko-polskoho seminaru. Eds. A. Bohutskyi, P. Hozhyk, M. Lanchont. Lviv, pp. 6–10.
5. Bradis Ye.M., Bachuryna H.F. 1969. Roslynnist URSR. Bolota. Kyiv: Naukova Dumka, 243 pp.
6. Brzeg A. 2005. Zespoły kserotermofilnych ziołorośli okraikowych z klasy Trifolio-Geranietea sanguinei Th. Müller 1962 w Polsce. Poznań: Bogucki Wydawnictwo Naukowe, 236 pp.
7. Didukh Ya.P. 2010. What is the future of our forests? Ukrainian Botanical Journal, (67)3: 321–343.
8. Didukh Ya.P. 2011. The ecological scales for the species of the Ukrainian flora and their use in synphytoindication. Kyiv: Phytosociocentre, 176 pp.
9. Didukh Ya.P. 2012. Fundamentals of bioindication. Kyiv: Naukova Dumka, 344 p.
10. Didukh Ya.P. 2023a. World of plants of Ukraine in aspect of the climate change. Kyiv: Naukova Dumka, 202 pp.
11. Didukh Ya.P. 2023b. Roslynnist Vyshnevoi hory ta yiyi okolyts. Kyiv; Chernivtsi: DrukArt, 136 pp.
12. Didukh Ya.P., Budzhak V.V. 2020. A program for automating the process of calculating indicator values of environmental factors: methodical recommendations. Chernivtsi: Yu. Fedkovych ChNU, 40 pp.
13. Didukh Ya.P., Plyuta P.H. 1994. Fitoindykatsiya ekolohichnyh faktoriv. Kyiv: Naukova Dumka, 280 pp.
14. Didukh Ya.P., Sheliah-Sosonko Yu.R. 2003. A geobotanical zoning of Ukraine and adjacent territories. Ukrainian Botanical Journal, 60(1): 6–17.
15. Dokturovskiy V.S. 1914. Predvaritelnyi otchet ob issledovaniyakh bolot Volynskoy gubernii v 1913 godu. Trudy Obshchestva issledovateley Volyni, 7: 161–179.
16. Dubyna D.V., Dziuba T.P., Iemelianova S.M., Bagrikova N.O., Borysova O.V., Borsukevych L.M., Vynokurov D.S., Gapon S.V., Gapon Yu.V., Davydov D.A., Dvoretskyi T.V., Didukh Ya.P., Zhmud O.I., Kozyr M.S., Konishchuk V.V., Kuzemko A.A., Pashkevych N.A., Ryff L.E., Solomakha V.A., Felbaba-Klushyna L.M., Fitsailo T.V., Chorna H.A., Chorney I.I., Shelyag-Sosonko Yu.R., Iakushenko D.M. 2019. Prodrome of the Vegetation of Ukraine. Eds D.V. Dubyna, T.P. Dziuba. Kyiv: Naukova Dumka, 782 pp.
17. Fijałkowski D., Izdebski K. 1957. Zbiorowiska stepowe na Wyżynie Lubelskiej. Annales Universitatis Mariae Curie-Sklodowska. Sectio C, 12(4): 167–200.
18. Fijałkowski D. 1966. Zbiorowiska roślinne lewobrzeżnej doliny Bugu w granicach województwa Lubelskiego. Annales Universitatis Mariae Curie-Sklodowska. Sectio C, 21: 247–312.
19. Fijałkowski D. 1967. Zbiorowiska ląkowo-pastwiskowe Lubelszczyzny. Zeszyty Problemowe Postępów Nauk Rolniczych, 74: 29–37.
20. Fijałkowski D. 1972. Stosunki geobotaniczne Lubelszczyzny. Lubelskie Towarzystwo Naukowe. Ossolineum, Wrocław, 285 pp.
21. Fijałkowski D. 1991. Zespoły roślinne Lubelszczyzny. Lublin: Wydawnictwo Uniwersytetu Marii Curie-Skłodowskiej, 303 pp.
22. Fijałkowski D. 1993. Lasy Lubelszczyzny. Lublin: LTN, 252 pp.
23. Fijałkowski D., Chojnacka-Fijalkowska E. 1990. Zbiorowiska z klas Phragmitea, Molinio-Arrhenatheretea i Scheuchzerio-Caricetea w makroregionie Lubelskim. Roczniki Nauk Rolniczych, Seria D. Monografie. Warszawa: Państwowe Wydawn. Naukowe, 217 pp.
24. Fiziko-geograficheskoe rayonirovanie Ukrainskoy SSR. 1968. Ed. V.P. Popov. Kyiv: Izdatelstvovo Kievskogo universiteta, 684 pp.
25. Barbarych A.I. 1977. Heobotanichne rayonuvannya Ukrainskoi RSR. Ed. A.I. Barbarych. Kyiv: Naukova Dumka, 330 pp.
26. Geographical encyclopedia of Ukraine. 1993. Vol. 3. Ed. O.M. Marynych. Kyiv: Ukrainska Radyanska Entsyklopediya im. M.P. Bazhana, 480 pp.
27. Hennekens S.M., Schaminée J.H.J. 2001. TURBOVEG, a comprehensive database management system for vegetation data. Journal of Vegetation Science, 12(4): 589–591. https://doi.org/10.2307/3237010
28. Herenchuk K.I. 1972. Pryroda Lvivskoi oblasti. Ed. K.I. Herenchuk. Lviv: Vydavnytstvo Lvivskoho universytetu, 151 pp.
29. Herenchuk K.I. 1975. Pryroda Volynskoi oblasti. Ed. K.I. Herenchuk. Lviv: Vyshcha shkola, 146 pp.
30. Herenchuk K.I. 1978. Pryroda Rovenskoi oblasti. Ed. K.I. Herenchuk. Lviv: Vydavnytstvo Lvivskoho universytetu, 148 pp.
31. Jahn A. 1956. Wyżyna Lubelska. Rzeźba i czwartorzęd. Prace Geograficzne. Vol. 7. Warszawa: Państwowe Wydawnictwo Naukowe, 448 pp.
32. Jutrzenka-Trzebiatowski A. 1995. Zboczowe lasy klonowo-lipowe Aceri-Tilietum Faber 1936 w Polsce Północno-Wschodniej. Monographiae Botanicae, 78: 1–78. https://doi.org/10.5586/mb.1995.002
33. Koczwara M. 1930. Geobotaniczne stosunki Wolynia. Rocznik Wołynski, 1: 7–56.
34. Kobendza R. 1933. Kilka uwag o roślinności plaskowzgorza Pelczańskiego na Wolyniu. Acta Societatis Botanicorum Poloniae, 10(2): 291–295.
35. Kucharczyk M. 1996a. Zespoły i zbiorowiska roślinne Kazimierskiego Parku Krajobrazowego. Cz. I. Zespoły łąkowe i pastwiskowe. Annales Universitatis Mariae Curie-Sklodowska. Sectio C, 51: 105–132.
36. Kucharczyk M. 1996b. Zespoły i zbiorowiska roślinne Kazimierskiego Parku Krajobrazowego. Cz. II. Zespoły wodne i szuwarowe. Annales Universitatis Mariae Curie-Sklodowska. Sectio C, 51: 133–183.
37. Kucharczyk M. 1998. Zespoły i zbiorowiska roślinne Kazimierskiego Parku Krajobrazowego. Cz. III. Zespoły leśne i zaroślowe. Annales Universitatis Mariae Curie-Sklodowska. Sectio C, 53: 139–165.
38. Kuzemko A.A. 2009. Roslynnist Ukrainy. Luchna roslynnist. Klas Molinio-Arrhenatheretea. Kyiv: Phytosociocentre, 376 pp.
39. Kuzmishyna I.I. 2008. Flora of Volhynian Upland, its antropic transformation and protection. Cand. Sci. Diss. Abstract. Kyiv, M.M. Gryshko National Botanic Garden, NAS of Ukraine, 34 pp.
40. Kuzmychov A.I. 1967a. Forests of the Volynian loess plateau. Ukrainian Botanical Journal, 24(2): 61–66.
41. Kuzmichov A.I. 1967b. Rastitelnost Volynskogo lessovogo plato i ee narodno–khozyaystvennoe znachenie. Avtoref. dis. ... kand. biol. nauk. Kyiv, 22 s.
42. Kuzmishyna I.I. 2008. Flora of Volhynian Upland, its antropic transformation and protection. Cand. Sci. Diss. Abstract. Kyiv, M.M. Gryshko National Botanic Garden NAS of Ukrainy, 34 pp.
43. Kwiatkowska A.J., Wyszomirski T. 1988. Decline of Potentillo albae-Quercetum phytocoenoses associated with the invasion of Carpinus betulus. Vegetatio, (75)1–2: 49–55. https://www.jstor.org/stable/20038275
44. Lentsevich S. 1959. Fizicheskaya geografiya Polshi. Moscow: Izdatelstvo inostrannoy literatury, 402 pp.
45. Lohvynenko I.P. 2009. Cladium mariscus na terytorii Volynskoi vysochyny. Naukovi zapysky Ternopilskoho natsionalnoho pedahohichnoho universytetu imeni Volodymyra Hnatyuka. Seriya: Biolohiya, 4(41): 59–63.
46. Logvynenko I.P. 2014. Rare and disappearing species of the flora of the Volyn Upland. Cand. Sci. Diss. Abstract. Kyiv, M.M. Gryshko National Botanic Garden NAS of Ukrainy, 20 pp.
47. Marinich A.M. 1978. Landshafty i fiziko-geograficheskoe rayonirovanie Ukrainskoy SSR. In: Atlas prirodnykh usloviy i estestvennykh resursov Ukrainskoy SSR. Moscow: GUGK pri SM SSSR, pp. 161–166.
48. Marinich A.M., Pashchenko V.M., Shishchenko P.G. 1985. The Nature of the Ukrainian SSR: Landscapes and physical geographical zoning. Kyiv: Naukova Dumka, 224 pp.
49. Matuszkiewicz W. 2002. Rrzewodnik do oznaczania zbiorowisk roslinnych Polski. Warszawa: PWN, 537 pp.
50. Melnyk V.Y., Baranskyi A.R. Mateychik V.I. 2006. The dynamics of the geographical range of Cladium mariscus (Cyperaceae) in Ukraine. Botanicheskiy Zhurnal, 91(4): 565–571.
51. Melnyk V.I., Parubok M.I., Savchuk R.V. 1999. Novi vidomosti pro stepovu roslynnist Volynskoi vysochyny. Ukrainian Phytosociological Collection. Series A. Phytosociology, 1–2(12–13): 30–33.
52. Mosyakin S.L., Fedoronchuk M.M. 1999. Vascular plants of Ukraine. A nomenclatural checklist. Kyiv, xxiii + 345 pp. https://doi.org/10.13140/2.1.2985.0409
53. Mucina L., Bültmann H., Dierßen K., Theurillat J.-P., Raus T., Čarni A., Šumberová K., Willner W., Dengler J., Gavilán García R., Chytrý M., Hájek M., Di Pietro R., Iakushenko D., Pallas J., Daniëls F.J.A., Bergmeier E., Santos Guerra A., Ermakov N., Valachovič M., Schaminée J.H.J., Lysenko T., Didukh Y.P., Pignatti S., Rodwell J.S., Capelo J., Weber H.E., Solomeshch A., Dimopoulos P., Aguiar C., Hennekens S.M., Tichý L. 2016. Vegetation of Europe: hierarchical floristic classification system of vascular plant, bryophyte, lichen, and algal communities. Applied Vegetation Science, 19(S1): 3–264. https://doi.org/10.1111/avsc.12257
54. Paczoski J. 1990. O formacyach roślinnych i o pochodzeniu fłory poleskiej. Pamięntnik fiziograficzny, 16: 1–156.
55. Panek J. 1931. Zespół wisienki stepowej (Prunetum fruticosae) i jego sukcesja. Rocznik Wołynski, 2: 403–424.
56. Pshevlotskyi M. 1999a. Erodovani grunty Sokalskoho pasma. Visnyk Lvivskoho universytetu. Seriya Heohrafichna, 24: 144–147.
57. Pshevlotskyi M. 1999b. Humusnyi stan erodovanykh gruntiv Sokalskoho pasma. Visnyk Lvivskoho universytetu. Seriya Heohrafichna, 25: 161–162.
58. Radziy V.F. 1998. Morfohenetychni osoblyvosti gruntiv Volynskoi vysochyny. Henezys, heohrafiya i ekolohiya gruntiv: Visnyk Lvivskoho universytetutu. Seriya Heohrafichna, 23: 233–238.
59. Sokolovskiy I.L. 1958. Lessovye porody Zapadnoy chasti USSR. Kyiv: Izdatelstvo AN USSR, 98 pp.
60. Szafer W. 1972. Szata roślinna Polski. Vols 1–2. 2^nd ed. Warszawa: PWN (Państwowe Wydawnictwo Naukowe), 346 pp.
61. Tarasyuk N., Fedas T. 2017. Ahroklimatychni resursy Volynskoi oblasti v umovakh suchasnoho klimatu. Youth Scientific Journal Lesya Ukrainka Eastern European National University, 9(358): 27–32.
62. Tichý L. 2002. JUICE, software for vegetation classification. Journal of Vegetation Science, 13: 451–453. https://doi.org/10.1111/j.1654-1103.2002.tb02069.x
63. Tsvetkov M.A. 1957. Izmenenie lesistosti Evropeyskoy Rossii s kontsa XVII stoletiya po 1914 god. Moscow: Izdatelstvo AN SSSR, 214 pp.
64. Uchbova karta gruntiv Zakhidnykh oblastey URSR. 1954. Ed. M.K. Krupskyi. Lviv: Kamenyar, 195 pp.
65. Vernarder N.B., Godlin M.M., Samburg G.N., Sporina S.A. 1951. Pochvy USSR. Kyiv; Kharkov: Gosselkhozizdat USSR, 326 pp.
66. Węgorek T. 1988. Zbiorowiska roślinności ciepłolubnej z udziałem krzewów i drzew na terenach erodowanych wyżyny Lubelskiej i Roztocza. Zeszyty problemowe postępów nauk rolniczych, 357: 135–152.
67. Zgłobicki W., Gawrysiak L., Baran-Zgłobicka B., Telecka M. 2016. Long-term forest cover changes, within an agricultural region, in relation to environmental variables, Lubelskie province, Eastern Poland. Environ Earth Sciences, 75: article number 1373. https://doi.org/10.1007/s12665-016-6195-z